Dr Mark Harvey

Senior Curator of Arachnology

Terrestrial Zoology

B.Sc., Ph.D.

"Dr Mark Harvey working at a microscope"


+61 (8) 9212 3737 mark.harvey@museum.wa.gov.au


  • Systematics of arachnids and myriapods, including their phylogenetic history and evolutionary relationships.
  • Taxonomy of arachnids and myriapods, and especially the documentation and description of new species.


  • Researches the taxonomy and systematics of arachnids and myriapods through the collections.
  • Develops the Museum’s collections through targeted fieldwork and collaborations with other organisations, including government, universities and industry.

Services (to other bodies):

  • Represents the Museum on committees and panels providing advice to government.


Exhibitions and Public Programs

  • Provides identifications and advice for environmental consultants, industry and government agencies.
  • Regularly presents talks and provides media coverage of work on arachnids and myriapods.
  • Co-editor of the Records of the Western Australian Museum; Associate Editor of the Journal of Arachnology and Invertebrate Systematics. Member of the editorial board of several other scholarly journals.
  • Co-supervisor to several post-graduate students at local and international universities.
  • Vice-President of the International Commission on Zoological Nomenclature: http://iczn.org/content/dr-mark-s-harvey

Selected Publications


Walter, D.E. and Harvey, M.S. (2009). Order Opilioacarida. In Krantz, G.W. and Walter, D.E. (ed.), Manual of Acarology, third edition: 104-106. Texas Tech University Press: Lubbock.

Harms, D. and Harvey, M.S. (2009). A review of the pirate spiders of Tasmania (Arachnida, Mimetidae, Australomimetus) with description of a new species. Journal of Arachnology 37: 188-205. http://www.americanarachnology.org/JoA_free/JoA_v37_n2/arac-37-02-188.pdf

Harms, D. and Harvey, M.S. (2009). Pirates of Australia: systematics and phylogeny of the Australian mimetid spiders (Araneae: Mimetidae: Mimetinae) with a description of the Western Australian fauna. Invertebrate Systematics 23: 231-280.

Korenko, S., Harvey, M.S. and Pekár, S. (2009). Stenochrus portoricensis new to the Czech Republic (Schizomida, Hubbardiidae). Arachnologische Mitteilungen 38: 1-3. http://journals.tubitak.gov.tr/zoology/issues/zoo-13-37-3/zoo-37-3-11-1210-9.pdf

Moir, M.L., Brennan, K.E.C. and Harvey, M.S. (2009). Diversity, endemism and species turnover of millipedes within the south-western Australia global biodiversity hotspot Journal of Biogeography 36: 1958-1971.

Jennings, J.T., Austin, A.D., Davies, K.A., Harvey, M.S., Hirst, D.B. and Taylor, G.S. (2009). Terrestrial invertebrates. In: Jennings, J.T. (ed.), Natural History of the Riverland and Murraylands: 306-333. Royal Society of South Australia Inc.: Adelaide.

Edward, K.L. and Harvey, M.S. (2009). A new species of Ischnothyreus (Araneae: Oonopidae) from monsoon rainforest of northern Australia. Records of the Western Australian Museum 25: 287-293. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_287-293_EDWARDetal.pdf

Rix, M.G., Roberts, J.D. and Harvey, M.S. (2009). The spider families Synotaxidae and Malkaridae (Arachnida: Araneae: Araneoidea) in Western Australia. Records of the Western Australian Museum 25: 295-304. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_295to304_RIXetal.pdf

Framenau, V.W., Main, B.Y., Harvey, M.S. and Waldock, J.M. (2009). Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae). Records of the Western Australian Museum 25: 309-314. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_309-314_FRAMENAUTELetal.pdf

Harvey, M.S. (2009). The first Australasian species of the halophilic pseudoscorpion genus Paraliochthonius (Pseudoscorpiones: Chthoniidae). Records of the Western Australian Museum 25: 329-344. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_329-344_HARVEY.pdf


Rix, M.G. and Harvey, M.S. (2010). The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level. Zookeys 36: 1-321. http://www.pensoft.net/journals/zookeys/article/306/the-spider-family-micropholcommatidae-arachnida-araneae-araneoidea-a-relimitation-and-revision-at-the-generic-level

Edward, K.L. and Harvey, M.S. (2010). A review of the Australian millipede genus Atelomastix (Diplopoda: Spirostreptida: Iulomorphidae). Zootaxa 2371: 1-63. http://www.mapress.com/zootaxa/2010/f/zt02371p063.pdf

Baehr, B.C., Harvey, M.S. and Smith, H.M. (2010). A review of the new endemic Australian goblin spider genus Cavisternum (Araneae: Oonopidae). American Museum Novitates 3684: 1-40. http://digitallibrary.amnh.org/dspace/handle/2246/6047

Rix, M.G. and Harvey, M.S. (2010). The first pararchaeid spider (Araneae: Pararchaeidae) from New Caledonia, with a discussion on spinneret spigots and egg sac morphology in Ozarchaea. Zootaxa 2414: 27-40. http://www.mapress.com/zootaxa/2010/f/z02414p040f.pdf

Burger, M., Harvey, M.S. and Stevens, N. (2010). A new species of blind subterranean Tetrablemma (Araneae: Tetrablemmidae) from Australia. Journal of Arachnology 38: 146-149. http://www.americanarachnology.org/JoA_free/JoA_v38_n1/arac-38-01-146.pdf

Harvey, M.S. and Muchmore, W.B. (2010). Two new cavernicolous species of the pseudoscorpion genus Cryptocreagris from Colorado (Pseudoscorpiones: Neobisiidae). Subterranean Biology 7: 55-64.

Framenau, V.W., Scharff, N. and Harvey, M.S. (2010). Systematics of the Australian orb-web spider genus Demadiana with comments on the generic classification of the Arkyinae (Araneae, Araneidae). Invertebrate Systematics 24: 139-171.

Durrant, B.J., Harvey, M.S., Framenau, V.W., Ott, R. and Waldock, J.M. (2010). Patterns in the composition of ground-dwelling spider communities in the Pilbara bioregion, Western Australia. Records of the Western Australian Museum, Supplement 78: 185-204.

Rix, M.G., Harvey, M.S. and Roberts, J.D. (2010). A revision of the textricellin spider genus Raveniella (Araneae: Araneoidea: Micropholcommatidae): exploring patterns of phylogeny and biogeography in an Australian biodiversity hotspot. Invertebrate Systematics 24: 209-237.

Harvey, M.S. (2010). A synopsis of the smaller arachnid orders of the Seychelles islands (Arachnida: Pseudoscorpiones, Schizomida, Amblypygi and Palpigradi). In: Gerlach, J. and Marusik, Y. (eds), Arachnida and Myriapoda of the Seychelles Islands: 331-340. Siri Scientific Press: Manchester.

Rahmadi, C., Harvey, M.S. and Kojima, J. (2010). Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island. Zootaxa 2612: 1-21. http://www.mapress.com/zootaxa/2010/f/z02612p021f.pdf

Harvey, M.S. and Šťáhlavský, F. (2010). A review of the pseudoscorpion genus Oreolpium (Pseudoscorpiones: Garypinidae), with remarks on the composition of the Garypinidae and on pseudoscorpions with bipolar distributions. Journal of Arachnology 38: 294-308. http://www.americanarachnology.org/JoA_free/JoA_v38_n2/arac-38-02-294.pdf

Harvey, M.S. (2010). Redescription of Geogarypus irrugatus from Sumatra (Pseudoscorpiones, Geogarypidae). Journal of Arachnology 38: 383-386. http://www.americanarachnology.org/JoA_free/JoA_v38_n2/arac-38-02-383.pdf

Baehr, B.C. and Harvey, M.S. (2010). Two new species of the endemic Australian goblin spider genus Cavisternum (Araneae: Oonopidae) from Queensland. Australian Entomologist 37: 171-177.

Volschenk, E.S., Burbidge, A.H., Durrant, B.J. and Harvey, M.S. (2010). Spatial distribution patterns of scorpions (Scorpiones) in the arid Pilbara region of Western Australia. Records of the Western Australian Museum, Supplement 78: 271-283.

Sirvid, P.J., Zhang, Z.-Q., Harvey, M.S., Rhode, B.E., Cook, D.R., Bartsch, I. and Staples, D.A. (2010). Subphylum Cheliceromorpha. Horseshoe crabs, sea spiders, and arachnids. In Gordon, D. (ed.), The New Zealand inventory of biodiversity: a Species 2000 Symposium review, vol. 1: 50-89. Canterbury University Press: Christchurch.


Harvey, M.S. (2011). Two new species of Synsphyronus (Pseudoscorpiones: Garypidae) from southern Western Australian granite landforms. Records of the Western Australian Museum 26: 11-22. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2010_26(1)_11to22_HARVEY.pdf

Wojcieszek, J.M., Harvey, M.S. and Rix, M.G. (2011). Optimised captive husbandry conditions for the Western Australian ‘Marri Millipede’ Antichiropus variabilis (Diplopoda: Polydesmida: Paradoxosomatidae), with notes on natural history. Records of the Western Australian Museum 26: 87-93. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2010_26(1)_87to93_WOJCIESZEKetal.pdf

Guzik, M.T., Austin, A.D., Cooper, S.J.B., Harvey, M.S., Humphreys, W.F., Bradford, T., Eberhard, S.M., King, R., Leys, R., Muirhead, K. and Tomlinson, M. (2011). Is the Australian subterranean fauna uniquely diverse? Invertebrate Systematics 24: 407-418.

Harvey, M.S. and Mahnert, V. (2011). Neobisium Chamberlin, 1930 (Arachnida, Pseudoscorpiones): proposed precedence over Blothrus Schiödte, 1847. Bulletin of Zoological Nomenclature 68: 47-53.

Crews, S.C. and Harvey, M.S. (2011). The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental region. Zookeys 99: 1-103.

Harvey, M.S. (2011). Cheiridium tetrophthalmum Daday, a new synonym of Larca lata (Hansen) (Pseudoscorpiones, Larcidae). Arachnologische Mitteilungen 41: 31-33.

Blick, T. and Harvey, M.S. (2011). Worldwide catalogues and species numbers of the arachnid orders (Arachnida). Arachnologische Mitteilungen 41: 41-43.

Harvey, M.S., Rix, M.G., Framenau, V.W., Hamilton, Z.R., Johnson, M.S., Teale, R.J., Humphreys, G. and Humphreys, W.F. (2011). Protecting the innocent: studying short-range endemic taxa enhances conservation outcomes. Invertebrate Systematics 25: 1-10.

Harvey, M.S. (2011). Notes on some Old World schizomids of the genera Ovozomus and Schizomus (Schizomida: Hubbardiidae). Records of the Western Australian Museum 26: 202-208. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2011_26(2)_202to208_HARVEY.pdf

Rix, M.G. and Harvey, M.S. (2011). Australian assassins, part I: A review of the Assassin Spiders (Araneae: Archaeidae) of mid-eastern Australia. Zookeys 123: 1-100. http://www.pensoft.net/journals/zookeys/article/1448/australian-assassins-part-i-a-review-of-the-assassin-spiders-araneae-archaeidae-of-mid-eastern-australia

Cooper, S.J.B., Harvey, M.S., Saint, K.M. and Main, B.Y. (2011). Deep phylogeographic structuring of populations of the trapdoor spider Moggridgea tingle (Migidae) from southwestern Australia: evidence for long-term refugia within refugia. Molecular Ecology 20: 3219-3236.

Rahmadi, C., Harvey, M.S. and Kojima, J. (2011).  The status of the whip spider subgenus Neocharon (Amblypygi: Charontidae) and the distribution of the genera Charon and Stygophrynus. Journal of Arachnology 39: 223-229. http://www.americanarachnology.org/JoA_free/JoA_v39_n2/arac-39-02-223.pdf

Harvey, M.S. (2011). Order Pseudoscorpiones de Geer, 1778. In: Zhang, Z.-Q. (ed.) Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 119-120.


Rix, M.G. and Harvey, M.S. (2012). Phylogeny and historical biogeography of ancient assassin spiders (Araneae: Archaeidae) in the Australian mesic zone: evidence for Miocene speciation within Tertiary refugia. Molecular Phylogenetics and Evolution 62: 375-396.

Platnick, N.I., Abrahim, N., Álvarez-Padilla, F., Andriamalala, D., Baehr, B.C., Baert, L., Bonaldo, A.B., Brescovit, A.D., Chousou-Polydouri, N., Dupérré, N., Eichenberger, B., Fannes, W., Gaublomme, E., Gillespie, R.G., Grismado, C.J., Griswold, C.E., Harvey, M.S., Henrard, A., Hormiga, G., Izquierdo, M.A., Jocqué, R., Kranz-Baltensperger, Y., Kropf, C., Ott, R., Ramirez, M.J., Raven, R.J., Rheims, C.A., Ruiz, G.R.S., Santos, A.J., Saucedo, A., Sierwald, P., Szüts, T., Ubick, D. and Wang, X.-P. (2012). Tarsal organ morphology and the phylogeny of goblin spiders (Araneae, Oonopidae), with notes on basal genera. American Museum Novitates 3736: 1-52. http://digitallibrary.amnh.org/dspace/handle/2246/6160

Rix, M.G. and Harvey, M.S. (2012). Australian assassins, part II: A review of the new assassin spider genus Zephyrarchaea (Araneae, Archaeidae) from southern Australia. Zookeys 191: 1-62. http://www.pensoft.net/journals/zookeys/article/3070/australian-assassins-part-ii-a-review-of-the-new-assassin-spider-genus-zephyrarchaea-araneae-archaeidae-from-southern-au

Wojcieszek, J.M., Austin, P., Harvey, M.S. and Simmons, L.W. (2012). Micro-CT scanning provides insight into the functional morphology of millipede genitalia. Journal of Zoology, London 287: 91-95.

Fontaine, B., Achterberg, K. van, Alonso-Zarazaga, M.A, Araujo, R., Asche, M., Aspöck, H., Aspöck, Audisio, P., Aukema, B., Bailly, N., Balsamo, M, Bank, R.A., Belfiore, C., Bogdanowicz, W., Boxshall, G., Burckhardt, D., Chylarecki, P., Deharveng, L., Dubois, A., Enghoff, H., Fochetti, R., Fontaine, C., Gargominy, O., Gomez Lopez, M.S., Goujet, D., Harvey, M.S., Heller, K.-G., Helsdingen, P. van, Hoch, H., Jong, Y. De, Karsholt, O., Los, W., Magowski, W, Massard, J.A., McInnes, S.J., Mendes, L.F., Mey, E., Michelsen, V., Minelli, A., Nieto Nafrıa, J.M., van Nieukerken, E.J., Pape, T., De Prins, W., Ramos, M., Ricci, C., Roselaar, C., Rota, E., Segers, H, Timm, T., van Tol, J. and Bouchet, P. (2012). New species in the Old World: Europe as a frontier in biodiversity exploration, a test bed for 21st century taxonomy. PloS One 7: e36881. http://www.plosone.org/article/info%3Adoi%2F10.1371%2Fjournal.pone.0036881

Harvey, M.S. (2012). A new species of Synsphyronus (Pseudoscorpiones: Garypidae) from Western Australia. Records of the Western Australian Museum 27: 55-61. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2012_27(1)_055to061_HARVEY.pdf

Baehr, B.C., Harvey, M.S., Burger, M. and Thoma, M. (2012). The new Australasian goblin spider genus Prethopalpus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History 763: 1-113. http://digitallibrary.amnh.org/dspace/handle/2246/6185

Volschenk, E.S., Harvey, M.S. and Prendini, L. (2012). A new species of Urodacus (Scorpiones: Urodacidae) from Western Australia. American Museum Novitates 3748: 1-18. http://digitallibrary.amnh.org/dspace/handle/2246/6181

Harvey, F.S.B., Framenau, V.W., Wojcieszek, J.M., Rix, M.G. and Harvey, M.S. (2012). Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia. Zootaxa 3383: 15-38. http://www.mapress.com/zootaxa/2012/f/z03383p038f.pdf

Sťáhlavský, F., Král, J., Harvey, M.S. and Haddad, C.R. (2012). The first cytogenetic characterisation of atemnids: pseudoscorpions with the highest chromosome numbers (Arachnida: Pseudoscorpiones). Cytogenetic and Genome Research 137: 22-30.

Rix, M.G. and Harvey, M.S. (2012). Australian assassins, part III: A review of the assassin spiders (Araneae, Archaeidae) of tropical north-eastern Queensland. Zookeys 218: 1-55. http://www.pensoft.net/journals/zookeys/article/3662/australian-assassins-part-iii-a-review-of-the-assassin-spiders-araneae-archaeidae-of-tropical-north-eastern-queensland

Harvey, M.S., Rahmadi, C., and Aland, K. (2012) The first record of whip spiders (Amblypygi) in Western Australia. Australasian Arachnology 84: 8-9.

Šťáhlavský, F., Boyer, S.L., Harvey, M.S. and Giribet, G. (2012). First cytogenetic study of a member of the harvestman family Pettalidae (Opiliones: Cyphophthalmi). Australian Journal of Entomology 51: 299-302.

Harvey, M.S., Ratnaweera, P.B. , Randeniya, P.V.  and Wijesinghe, M.R. (2012). A new species of the pseudoscorpion genus Megachernes (Pseudoscorpiones: Chernetidae) associated with a threatened Sri Lankan rainforest rodent, with a review of host associations of MegachernesJournal of Natural History 46: 2519-2535.


Car, C.A. and Harvey, M.S. (2013). A review of the Western Australian keeled millipede genus Boreohesperus (Diplopoda: Polydesmida: Paradoxosomatidae). Zookeys 290: 1–19. http://www.pensoft.net/journals/zookeys/article/5114/a-review-of-the-western-australian-keeled-millipede-genus-boreohesperus-diplopoda-polydesmida-paradoxosomatidae-

Zawierucha, K., Szymkowiak, P., Dabert, M. and Harvey, M.S. (2013). First record of the schizomid Stenochrus portoricensis (Schizomida: Hubbardiidae) in Poland, with DNA barcode data. Turkish Journal of Arachnology 37: 357–361. http://journals.tubitak.gov.tr/zoology/issues/zoo-13-37-3/zoo-37-3-11-1210-9.pdf

Baehr, B.C. and Harvey, M.S. (2013). The first goblin spiders of the genus Camptoscaphiella (Araneae: Oonopidae) from New Caledonia. Australian Journal of Entomology 52: 144–150.

Harms, D. and Harvey, M.S. (2013). Review of the cave-dwelling species of Pseudotyrannochthonius Beier (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae) from mainland Australia, with description of two troglobitic species. Australian Journal of Entomology 52: 129–143.

Monod, L., Harvey, M.S. and Prendini, L. (2013). Stenotopic Hormurus Thorell, 1876 scorpions from the monsoon ecosystems of northern Australia, with a discussion on the evolution of burrowing behaviour in Hormuridae Laurie, 1896. Revue Suisse de Zoologie 120: 281–346.

Harvey, M.S. (2013). Order Pseudoscorpiones. Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703: 34–35. http://biotaxa.org/Zootaxa/article/view/zootaxa.3703.1.8/4262

Girişgin, A.O., Girişgin, O. and Harvey, M.S. (2013). The first record of pseudoscorpions in honeybee hives in Turkey. Turkish Journal of Veterinary and Animal Sciences 37: 608–610. http://journals.tubitak.gov.tr/veterinary/issues/vet-13-37-5/vet-37-5-21-1207-7.pdf

Nassirkhani, M. and Harvey, M.S. (2013). Description of the postlarval stages of Dactylochelifer gracilis Beier, Pseudoscorpiones: Cheliferidae. International Journal of Zoology 2013(782638): 1–15. http://www.hindawi.com/journals/ijz/2013/782638/

Baehr, B.C., Harvey, M.S., Smith, H.M. and Ott, R. (2013). The goblin spider genus Opopaea in Australia and the Pacific islands (Araneae: Oonopidae). Memoirs of the Queensland Museum – Nature58: 107–338.

Harvey, M.S. and Baehr, B. (2013). Two new species of the goblin spider genus Cavisternum from tropical Australia (Araneae: Oonopidae). Memoirs of the Queensland Museum - Nature 58: 359–366.

Harvey, M.S. and Muchmore, W.B. (2013). The systematics of the pseudoscorpion family Ideoroncidae (Pseudoscorpiones, Neobisioidea) in the New World. Journal of Arachnology 41: 229–290.

Gunawardene, N.R., Taylor, C.K., Majer, J.D. and Harvey, M.S. (eds) (2013). The terrestrial invertebrate fauna of Barrow Island, Western Australia. Records of the Western Australian Museum, Supplement. Western Australian Museum: Perth. http://museum.wa.gov.au/store/museum-books/supplements/supplement-83-terrestrial-invertebrate-fauna-barrow-island

Humphreys, G., Alexander, J., Harvey, M.S. and Humphreys, W.F. (2013). The subterranean fauna of Barrow Island, north-western Australia: 10 years on. Records of the Western Australian Museum, Supplement 83: 145–158.

Car, C.A., Short, M., Huynh, C. and Harvey, M.S. (2013). The millipedes of Barrow Island, Western Australia (Diplopoda). Records of the Western Australian Museum, Supplement 83: 209–219.

Majer, J.D., Gunawardene, N.R., Taylor, C.K. and Harvey, M.S. (2013). A last word. Records of the Western Australian Museum, Supplement 83: 405–406.

Car, C.A., Wojcieszek, J.M. and Harvey, M.S. (2013). The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae) Part 1: Redefinition of the genus and redescriptions of existing species. Records of the Western Australian Museum 28: 83–118. http://museum.wa.gov.au/sites/default/files/WAMRecords_2013_28(2)_83to118_CARetal.pdf


Harvey, M.S. (2014). A review and redescription of the cosmopolitan pseudoscorpion Chelifer cancroides (Pseudoscorpiones: Cheliferidae). Journal of Arachnology 42: 86–104

Harvey, M.S. and Du Preez, G. (2014). A new troglobitic ideoroncid pseudoscorpion (Pseudoscorpiones: Ideoroncidae) from southern Africa. Journal of Arachnology 42: 105–110.

Miglio, L.T., Harms, D., Framenau, V.W. and Harvey, M.S. (2014). Four new mouse spider species (Araneae, Mygalomorphae, Actinopodidae, Missulena) from Western Australia. ZooKeys 410: 121–148.

Edward, K.L. and Harvey, M.S. (2014). Australian goblin spiders of the genus Ischnothyreus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History 389: 1–144.

Car, C.A. and Harvey, M.S. (2014). The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 2: species of the Great Western Woodlands region of Western Australia. Records of the Western Australian Museum 29: 20–77.

Harvey, M.S. (2014). Arachnida (Arthropoda: Chelicerata) of Western Australia: overview and prospects. Journal of the Royal Society of Western Australia 97: 57–64.

Harrison, S.E., Guzik, M.T., Harvey, M.S. and Austin, A.D. (2014). Molecular phylogenetic analysis of Western Australian troglobitic chthoniid pseudoscorpions (Pseudoscorpiones : Chthoniidae) points to multiple independent subterranean clades. Invertebrate Systematics 28: 386–400.

Castalanelli, M.A., Teale, R., Rix, M.G., Kennington, J.W. and Harvey, M.S. (2014). Barcoding of mygalomorph spiders (Araneae: Mygalomorphae) in the Pilbara bioregion of Western Australia reveals a hgihly diverse biota. Invertebrate Systematics 28: 375–385.

Giribet, G., McIntyre, E., Christian, E., Espinasa, L., Ferreira, R.L., Francke, O.F., Harvey, M.S., Isaia, M., Kovác, L., McCutchen, L., Souza, M.F.V.R. and Zagmajster, M. (2014). The first phylogenetic analysis of Palpigradi (Arachnida)—the most enigmatic arthropod order. Invertebrate Systematics 28: 350–360.

Alexander, J.B., Burger, M.A.A. and Harvey, M.S. (2014). A new species of troglobitic Anatemnus (Pseudoscorpiones: Atemnidae) from the Pilbara bioregion of Australia. Records of the Western Australian Museum 29: 141–148.

Harvey, M.S. and Wynne, J.J. (2014). Troglomorphic pseudoscorpions (Arachnida: Pseudoscorpiones) of northern Arizona, with the description of two new short-range endemic species. Journal of Arachnology 42: 205–219.


Moir, M.L., Comer, S., and Harvey, M.S. (2015). The benefits of multi-actor invertebrate management. Decision Point 87: 6–7.

Rix, M.G., Edwards, D.L., Byrne, M., Harvey, M.S., Joseph, L. and Roberts, J.D. (2015). Biogeography and speciation of terrestrial fauna in the south-western Australian biodiversity hotspot. Biological Reviews 90: 762–793.

Harvey, M.S. and Mahnert, V. (2015). A new species of Sphaerowithius (Pseudoscorpiones, Withiidae) from Namibia. African Invertebrates 90: 491–497.

Harvey, M.S., Lopes, P.C., Goldsmith, G.R., Halajian, A., Hillyer, M. and Huey, J.A. (2015). A novel symbiotic relationship between sociable weaver birds (Philetairus socius) and a new cheliferid pseudoscorpion (Pseudoscorpiones: Cheliferidae) in southern Africa. Invertebrate Systematics 29: 444–456.

Harvey, M.S. (2015). A review of the taxonomy and biology of pseudoscorpions of Nannowithius and Termitowithius (Pseudoscorpiones, Withiidae), inquilines of social insects. Journal of Arachnology 43: 342–352.

Harvey, M.S. (2015). Revised diagnoses for the pseudoscorpion genera Metawithius and Microwithius, with the description of a new Australian genus, and notes on Withius (Pseudoscorpiones, Withiidae). Journal of Arachnology 43: 353–370.

Ott, R., Von Beeren, C. and Harvey, M.S. (2015). Sicariomorpha, a new myrmecophilous goblin spider genus (Araneae, Oonopidae) associated with Asian army ants. American Museum Novitates 3843: 1–14.

Novák, J. and Harvey, M.S. (2015). The identity of pseudoscorpions of the genus Diplotemnus (Pseudoscorpiones: Atemnidae) from Europe and Asia. North-Western Journal of Zoology 11: 316–323.

Abrams, K.M. and Harvey, M.S. (2015). A new troglobitic schizomid (Hubbardiidae: Paradraculoides) from the Pilbara region, Western Australia. Records of the Western Australian Museum 30: 132–136.

Harvey, M.S., Abrams, K.M. and Burger, M.A.A. (2015). A new species of the pseudoscorpion genus Synsphyronus (Pseudoscorpiones: Garypidae) from Barrow Island, Western Australia. Records of the Western Australian Museum 30: 137–143.

Finlayson, G.R., Madani, G., Dennis, G., and Harvey, M. (2015). First reported observation of phoresy of pseudoscorpions on an endemic New Zealand mammal, the lesser short-tailed bat, Mystacina tuberculata. New Zealand Journal of Zoology 42: 298–301.

Harvey, M.S., Main, B.Y., Rix, M.G. and Cooper, S.J.B. (2015). Refugia within refugia: in situ speciation and conservation of threatened Bertmainius (Araneae: Migidae), a new genus of relictual trapdoor spiders endemic to the mesic zone of south-western Australia. Invertebrate Systematics 29: 511–553.


Harvey, M.S., Huey, J.A., Hillyer, M.A., McIntyre, E. and Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics 30: 75–85.

Downes, M.F. and Harvey, M.S. (2016). The spider Oecobius concinnus Simon (Araneae: Oecobiidae) in nests of the ant Polyrhachis australis Mayr (Hymenoptera: Formicidae) in Townsville, Queensland: a new distribution record. Australian Entomologist 43: 31–34.

Harrison, S.E., Rix, M.G., Harvey, M.S. and Austin, A.D. (2016). An African mygalomorph lineage in temperate Australia: the trapdoor spider genus Moggridgea (Araneae: Migidae) on Kangaroo Island, South Australia. Austral Entomology 55: 208–216.

Harvey, M.S., Andrade, R. and Pinto-da-Rocha, R. (2016). The first New World species of the pseudoscorpion family Feaellidae (Pseudoscorpiones: Feaelloidea), from the Brazilian Atlantic Forest. Journal of Arachnology 44: 237–234.

Giribet, G., Boyer, S.L., Baker, C., Fernández, R., Sharma, P., De Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. and Griswold, C.E. (2016). A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) with biogeographic and taxonomic implications. Zoological Journal of the Linnean Society 178: 523–545.

Harvey, M.S., Abrams, K.M., Beavis, A.S., Hillyer, M.J. and Huey, J.A. (2016). Pseudoscorpions of the family Feaellidae (Pseudoscorpiones: Feaelloidea) from the Pilbara region of Western Australia show extreme short-range endemism. Invertebrate Systematics 30: 491–508.

Harvey, M.S. and Bosmans, R. (2016). Order Pseudoscorpiones, pseudoscorpions. In: D. J. Sparrow and E. John, An introduction to the wildlife of Cyprus: 521–524. (Terra Cypria: Lemesos).

Cosgrove, J.G., Agnarsson, I., Harvey, M.S. and Binford, G.J. (2016). Pseudoscorpion diversity and distribution in the West Indies: sequence data confirm single island endemism for some clades, but not others. Journal of Arachnology 44: 257–271.

Harvey, M.S. (2016). The systematics of the pseudoscorpion family Ideoroncidae (Pseudoscorpiones, Neobisioidea) in the Asian region. Journal of Arachnology 44: 272–329.


Harvey, M.S., Rix, M.G., Harms, D., Giribet, G., Vink, C.J. and Walter, D.E. (2017). The biogeography of Australasian arachnids. In: M. Ebach (ed.), Handbook on Australasian Biogeography. (CRC Press / Taylor and Francis: Boca Raton).

Rix, M.G., Cooper, S.J.B., Meusemann, K., Klopfstein, S., Harrison, S.E., Harvey, M.S. and Austin, A.D. (2017). Post-Eocene climate change across continental Australia and the diversification of Australasian spiny trapdoor spiders (Idiopidae: Arbanitinae) Molecular Phylogenetics and Evolution 109: 302–320.

Rix, M.G., Huey, J.A., Main, B.Y., Waldock, J.M., Harrison, S.E., Comer, S., Austin, A.D. and Harvey, M.S. (2017). Where have all the spiders gone? Highlighting the decline of a poorly known invertebrate fauna in the agricultural and arid zones of southern Australia. Austral Entomology 56: 14–22.

Gao, Z., Zhang, F. and Harvey, M.S. (2017). A modified definition of the genus Haplochernes (Pseudoscorpiones: Chernetidae), with a new species from Hainan Island. Journal of Arachnology 45: 112–122.

Car, C.A. and Harvey, M.S. (2017). New species of Boreohesperus (Diplopoda, Polydesmida, Paradoxosomatidae) from north-western Australia. European Journal of Taxonomy 320: 1–11.

Smith, H.M., Harvey, M.S., Agnarsson, I. and Anderson, G.J. (2017). Notes on the ant-mimic genus Anatea Berland (Araneae: Theridiidae) and two new species from tropical Australia. Records of the Australian Museum 69: 1–13.

Castalanelli, M.A., Huey, J.A., Hillyer, M.J. and Harvey, M.S. (2017). Molecular and morphological evidence for a new genus of small trapdoor spiders from arid Western Australia (Araneae: Mygalomorphae: Nemesiidae: Anaminae). Invertebrate Systematics 31: 492–505.

Harrison, S.E., Harvey, M.S., Cooper, S.J.B., Austin, A.D. and Rix, M.G. (2017). Across the Indian Ocean: A remarkable example of trans-oceanic dispersal in an austral mygalomorph spider. PLoS One. 12(8) e0180139: 1–16.

Rix, M.G., Raven, R.J., Main, B.Y., Harrison, S.E., Austin, A.D., Cooper, S.J.B. and Harvey, M.S. (2017). The Australasian spiny trapdoor spiders of the family Idiopidae (Mygalomorphae: Arbanitinae): a relimitation and revision at the generic level. Invertebrate Systematics 31: 566–634.

Clouse, R.M., Branstetter, M.G., Buenavente, P., Crowley, L.M., Czekanski-Moir, J., Emmanuel M. General, D., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Mohagan, D.P., Sharma, P.P. and Wheeler, W.C. (2017). First global molecular phylogeny and biogeographical analysis of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44: 2660–2672.

Rix, M.G., Bain, K., Main, B.Y., Raven, R.J., Austin, A.D., Cooper, S.J.B. and Harvey, M.S. (2017). The  spiny trapdoor spiders of the genus Cataxia (Mygalomorphae: Idiopidae) from south-western Australia: documenting a threatened fauna in a sky-island landscape. Journal of Arachnology 45: 395–423.

Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. and Giribet, G. (2018). A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122: 37–45.

Schmidt, A.R., Kaulfuss, U., Bannister, J.M., Baranov, V., Beimforde, C., Bleile, N., Borkent, A., Busch, A., Conran, J.G., Engel, M.S., Harvey, M., Kennedy, E.M., Kerr, P.H., Kettunen, E., Kiecksee, A.P., Lengeling, F., Lindqvist, J.K., Maraun, M., Mildenhall, D.C., Perrichot, V., Rikkinen, J., Sadowski, E.-M., Seyfullah, L.J., Stebner, F., Szwedo, J., Ulbrich, P. and Lee, D.E. (2018). Amber inclusions from New Zealand. Gondwana Research 56: 135–146.

Stephen, C.D.R. and Harvey, M.S. (2018). William B. Muchmore (1920–2017): his taxonomic contributions and complete bibliography. Journal of Cave and Karst Studies 80: 39–48.

Thomson, S.A., Pyle, R.L., Ahyong, S., Alonso-Zarazaga, M., Ammirati, J., Araya, J.F., Ascher, J.S., Audisio, T.L., Azevedo-Santos, V.M., Bailly, N., Baker, W.J., Balke, M., Barclay, M.V.L., Barrett, R.L., Benine, R.C., Bickerstaff, J.R.M., Bouchard, P., Bour, R., Bourgoin, T., Boyko, C.B., Breure, A.S.H., Brothers, D.J., Byng, J.W., Campbell, D., P.Ceriaco, L.M., Cernak, I., Cerretti, P., Chang, C.-H., Cho, S., Copus, J.M., Costello, M.J., Cseh, A., Csuzdi, C., Culham, A., D’Elia, G., d’Acoz, C.d.U., Daneliya, M.E., Dekker, R., Dickinson, E.C., Dickinson, T.A., Dijk, P.P.v., Dijkstra, K.-D.B., Dima, B., Dmitriev, D.A., Duistermaat, L., Dumbacher, J.P., Eiserhardt, W.L., Ekrem, T., Evenhuis, N.L., Faille, A., Fernandez-Triana, J.L., Fiesler, E., Fishbein, M., Fordham, B.G., Freitas, A.V.L., Friol, N.R., Fritz, U., Froslev, T., Funk, V.A., Gaimari, S.D., Garbino, G.S.T., S.Garraffoni, A.R., Geml, J., Gill, A.C., Gray, A., Grazziotin, F.G., Greenslade, P., Gutierrez, E.E., Harvey, M.S., Hazevoet, C.J., He, K., He, X., Helfer, S., Helgen, K.M., Heteren, A.H.v., Garcia, F.H., Holstein, N., Horvath, M.K., Hovenkamp, P.H., Hwang, W.S., Hyvonen, J., Islam, M.B., Iverson, J.B., Ivie, M.A., Jaafar, Z., Jackson, M.D., Jayat, J.P., Johnson, N.F., Kaiser, H., Klitgard, B.B., Knapp, D.G., Kojima, J.-i., Koljalg, U., Kontschan, J., Krell, F.-T., Krisai-Greilhuber, I., Kullander, S., Latella, L., Lattke, J.E., Lencioni, V., Lewis, G.P., Lhano, M.G., Lujan, N.K., Luksenburg, J.A., Mariaux, J., Marinho-Filho, J., Marshall, C.J., Mate, J.F., McDonough, M.M., Michel, E., Miranda, V.F.O., Mitroiu, M.-D., Molinari, J., Monks, S., Moore, A.J., Moratelli, R., Muranyi, D., Nakano, T., Nikolaeva, S., Noyes, J., Ohl, M., Oleas, N.H., Orrell, T., Pall-Gergely, B., Pape, T., Papp, V., Parenti, L.R., Patterson, D., Pavlinov, I.Y., Pine, R.H., Poczai, P., Prado, J., Prathapan, D., Rabeler, R.K., Randall, J.E., Rheindt, F.E., Rhodin, A.G.J., Rodriguez, S.M., Rogers, D.C., Roque, F.d.O., Rowe, K.C., Ruedas, L.A., Salazar-Bravo, J., Salvador, R.B., Sangster, G., Sarmiento, C.E., S.Schigel, D., Schmidt, S., Schueler, F.W., Segers, H., Snow, N., Souza-Dias, P.G.B., Stals, R., Stenroos, S., Stone, R.D., Sturm, C.F., Štys, P., Teta, P., Thomas, D.C., Timm, R.M., Tindall, B.J., Todd, J.A., Triebel, D., Valdecasas, A.G., Vizzini, A., Vorontsova, M.S., Vos, J.M.d., Wagner, P., Watling, L., Weakley, A., Welter-Schultes, F., Whitmore, D., Wilding, N., Will, K., Williams, J., Wilson, K., Winston, J.E., Wuster, W., Yanega, D., Yeates, D.K., Zaher, H., Zhang, G., Zhang, Z.-Q. and Zhou, H.-Z. (2018). Taxonomy based on science is necessary for global conservation. PloS Biology 16 (3): e2005075.

Novák, J. and Harvey, M.S. (2018). New species and records of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from India, Sri Lanka and New Guinea. Zootaxa 4394: 417–427.

Rix, M.G., Main, B.Y., Raven, R.J. and Harvey, M.S. (2018). Systematics of the spiny trapdoor spiders of the genus Eucanippe (Mygalomorphae: Idiopidae: Aganippini) from south-western Australia: documenting a poorly-known lineage from Australia's biodiversity hotspot. Journal of Arachnology 46: 133–154.

Harvey, M.S., Cosgrove, J.G., Harms, D., Selden, P.A., Shih, C. and Wang, C.-C. (2018). The oldest chthonioid pseudoscorpion (Arachnida: Pseudoscorpiones: Chthonioidea: Chthoniidae): a new genus and species from mid-Cretaceous Burmese amber. Zoologischer Anzeiger 273: 102–111.

Harvey, M.S. (2018). Two new species of the pseudoscorpion genus Cybella (Pseudoscorpiones: Feaellidae) from Malaysian caves. Zoologischer Anzeiger 273: 124–132.

Rix, M.G., Huey, J.A., Cooper, S.J.B., Austin, A.D. and Harvey, M.S. (2018). Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia. ZooKeys 756: 1–121.