Mark Harvey

Senior Curator of Arachnology

Terrestrial Zoology

B.Sc., Ph.D.

Mark Harvey working at a microscope

Contact

+61 (8) 9212 3737 mark.harvey@museum.wa.gov.au

Research


  • Systematics of arachnids and myriapods, including their phylogenetic history and evolutionary relationships.
  • Taxonomy of arachnids and myriapods, and especially the documentation and description of new species.

Collections:

  • Researches the taxonomy and systematics of arachnids and myriapods through the collections.
  • Develops the Museum’s collections through targeted fieldwork and collaborations with other organisations, including government, universities and industry.

Services (to other bodies):

  • Represents the Museum on committees and panels providing advice to government.

Websites


  • Pseudoscorpions of the World: http://www.museum.wa.gov.au/catalogues/pseudoscorpions/
  • WAMinals: http://museum.wa.gov.au/catalogues/waminals
  • Arachnids of the World (BETA): http://museum.wa.gov.au/catalogues-beta/

Exhibitions and Public Programs


  • Provides identifications and advice for environmental consultants, industry and government agencies.
  • Regularly presents talks and provides media coverage of work on arachnids and myriapods.
  • Co-editor of the Records of the Western Australian Museum; Associate Editor of the Journal of Arachnology and Invertebrate Systematics. Member of the editorial board of several other scholarly journals.
  • Co-supervisor to several post-graduate students at local and international universities.

Selected Publications

Mark Harvey's Publications 2009 - 2013

2009

Walter, D.E. and Harvey, M.S. (2009). Order Opilioacarida. In Krantz, G.W. and Walter, D.E. (ed.), Manual of Acarology, third edition: 104-106. Texas Tech University Press: Lubbock.

Harms, D. and Harvey, M.S. (2009). A review of the pirate spiders of Tasmania (Arachnida, Mimetidae, Australomimetus) with description of a new species. Journal of Arachnology 37: 188-205. http://www.americanarachnology.org/JoA_free/JoA_v37_n2/arac-37-02-188.pdf

Harms, D. and Harvey, M.S. (2009). Pirates of Australia: systematics and phylogeny of the Australian mimetid spiders (Araneae: Mimetidae: Mimetinae) with a description of the Western Australian fauna. Invertebrate Systematics 23: 231-280.

Korenko, S., Harvey, M.S. and Pekár, S. (2009). Stenochrus portoricensis new to the Czech Republic (Schizomida, Hubbardiidae). Arachnologische Mitteilungen 38: 1-3. http://journals.tubitak.gov.tr/zoology/issues/zoo-13-37-3/zoo-37-3-11-1210-9.pdf

Moir, M.L., Brennan, K.E.C. and Harvey, M.S. (2009). Diversity, endemism and species turnover of millipedes within the south-western Australia global biodiversity hotspot Journal of Biogeography 36: 1958-1971.

Jennings, J.T., Austin, A.D., Davies, K.A., Harvey, M.S., Hirst, D.B. and Taylor, G.S. (2009). Terrestrial invertebrates. In: Jennings, J.T. (ed.), Natural History of the Riverland and Murraylands: 306-333. Royal Society of South Australia Inc.: Adelaide.

Edward, K.L. and Harvey, M.S. (2009). A new species of Ischnothyreus (Araneae: Oonopidae) from monsoon rainforest of northern Australia. Records of the Western Australian Museum 25: 287-293. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_287-293_EDWARDetal.pdf

Rix, M.G., Roberts, J.D. and Harvey, M.S. (2009). The spider families Synotaxidae and Malkaridae (Arachnida: Araneae: Araneoidea) in Western Australia. Records of the Western Australian Museum 25: 295-304. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_295to304_RIXetal.pdf

Framenau, V.W., Main, B.Y., Harvey, M.S. and Waldock, J.M. (2009). Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae). Records of the Western Australian Museum 25: 309-314. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_309-314_FRAMENAUTELetal.pdf

Harvey, M.S. (2009). The first Australasian species of the halophilic pseudoscorpion genus Paraliochthonius (Pseudoscorpiones: Chthoniidae). Records of the Western Australian Museum 25: 329-344. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_25(3)_2009_329-344_HARVEY.pdf

2010

Rix, M.G. and Harvey, M.S. (2010). The spider family Micropholcommatidae (Arachnida: Araneae: Araneoidea): a relimitation and revision at the generic level. Zookeys 36: 1-321. http://www.pensoft.net/journals/zookeys/article/306/the-spider-family-micropholcommatidae-arachnida-araneae-araneoidea-a-relimitation-and-revision-at-the-generic-level

Edward, K.L. and Harvey, M.S. (2010). A review of the Australian millipede genus Atelomastix (Diplopoda: Spirostreptida: Iulomorphidae). Zootaxa 2371: 1-63. http://www.mapress.com/zootaxa/2010/f/zt02371p063.pdf

Baehr, B.C., Harvey, M.S. and Smith, H.M. (2010). A review of the new endemic Australian goblin spider genus Cavisternum (Araneae: Oonopidae). American Museum Novitates 3684: 1-40. http://digitallibrary.amnh.org/dspace/handle/2246/6047

Rix, M.G. and Harvey, M.S. (2010). The first pararchaeid spider (Araneae: Pararchaeidae) from New Caledonia, with a discussion on spinneret spigots and egg sac morphology in Ozarchaea. Zootaxa 2414: 27-40. http://www.mapress.com/zootaxa/2010/f/z02414p040f.pdf

Burger, M., Harvey, M.S. and Stevens, N. (2010). A new species of blind subterranean Tetrablemma (Araneae: Tetrablemmidae) from Australia. Journal of Arachnology 38: 146-149. http://www.americanarachnology.org/JoA_free/JoA_v38_n1/arac-38-01-146.pdf

Harvey, M.S. and Muchmore, W.B. (2010). Two new cavernicolous species of the pseudoscorpion genus Cryptocreagris from Colorado (Pseudoscorpiones: Neobisiidae). Subterranean Biology 7: 55-64.

Framenau, V.W., Scharff, N. and Harvey, M.S. (2010). Systematics of the Australian orb-web spider genus Demadiana with comments on the generic classification of the Arkyinae (Araneae, Araneidae). Invertebrate Systematics 24: 139-171.

Durrant, B.J., Harvey, M.S., Framenau, V.W., Ott, R. and Waldock, J.M. (2010). Patterns in the composition of ground-dwelling spider communities in the Pilbara bioregion, Western Australia. Records of the Western Australian Museum, Supplement 78: 185-204.

Rix, M.G., Harvey, M.S. and Roberts, J.D. (2010). A revision of the textricellin spider genus Raveniella (Araneae: Araneoidea: Micropholcommatidae): exploring patterns of phylogeny and biogeography in an Australian biodiversity hotspot. Invertebrate Systematics 24: 209-237.

Harvey, M.S. (2010). A synopsis of the smaller arachnid orders of the Seychelles islands (Arachnida: Pseudoscorpiones, Schizomida, Amblypygi and Palpigradi). In: Gerlach, J. and Marusik, Y. (eds), Arachnida and Myriapoda of the Seychelles Islands: 331-340. Siri Scientific Press: Manchester.

Rahmadi, C., Harvey, M.S. and Kojima, J. (2010). Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island. Zootaxa 2612: 1-21. http://www.mapress.com/zootaxa/2010/f/z02612p021f.pdf

Harvey, M.S. and Šťáhlavský, F. (2010). A review of the pseudoscorpion genus Oreolpium (Pseudoscorpiones: Garypinidae), with remarks on the composition of the Garypinidae and on pseudoscorpions with bipolar distributions. Journal of Arachnology 38: 294-308. http://www.americanarachnology.org/JoA_free/JoA_v38_n2/arac-38-02-294.pdf

Harvey, M.S. (2010). Redescription of Geogarypus irrugatus from Sumatra (Pseudoscorpiones, Geogarypidae). Journal of Arachnology 38: 383-386. http://www.americanarachnology.org/JoA_free/JoA_v38_n2/arac-38-02-383.pdf

Baehr, B.C. and Harvey, M.S. (2010). Two new species of the endemic Australian goblin spider genus Cavisternum (Araneae: Oonopidae) from Queensland. Australian Entomologist 37: 171-177.

Volschenk, E.S., Burbidge, A.H., Durrant, B.J. and Harvey, M.S. (2010). Spatial distribution patterns of scorpions (Scorpiones) in the arid Pilbara region of Western Australia. Records of the Western Australian Museum, Supplement 78: 271-283.

Sirvid, P.J., Zhang, Z.-Q., Harvey, M.S., Rhode, B.E., Cook, D.R., Bartsch, I. and Staples, D.A. (2010). Subphylum Cheliceromorpha. Horseshoe crabs, sea spiders, and arachnids. In Gordon, D. (ed.), The New Zealand inventory of biodiversity: a Species 2000 Symposium review, vol. 1: 50-89. Canterbury University Press: Christchurch.

2011

Harvey, M.S. (2011). Two new species of Synsphyronus (Pseudoscorpiones: Garypidae) from southern Western Australian granite landforms. Records of the Western Australian Museum 26: 11-22. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2010_26(1)_11to22_HARVEY.pdf

Wojcieszek, J.M., Harvey, M.S. and Rix, M.G. (2011). Optimised captive husbandry conditions for the Western Australian ‘Marri Millipede’ Antichiropus variabilis (Diplopoda: Polydesmida: Paradoxosomatidae), with notes on natural history. Records of the Western Australian Museum 26: 87-93. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2010_26(1)_87to93_WOJCIESZEKetal.pdf

Guzik, M.T., Austin, A.D., Cooper, S.J.B., Harvey, M.S., Humphreys, W.F., Bradford, T., Eberhard, S.M., King, R., Leys, R., Muirhead, K. and Tomlinson, M. (2011). Is the Australian subterranean fauna uniquely diverse? Invertebrate Systematics 24: 407-418.

Harvey, M.S. and Mahnert, V. (2011). Neobisium Chamberlin, 1930 (Arachnida, Pseudoscorpiones): proposed precedence over Blothrus Schiödte, 1847. Bulletin of Zoological Nomenclature 68: 47-53.

Crews, S.C. and Harvey, M.S. (2011). The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental region. Zookeys 99: 1-103.

Harvey, M.S. (2011). Cheiridium tetrophthalmum Daday, a new synonym of Larca lata (Hansen) (Pseudoscorpiones, Larcidae). Arachnologische Mitteilungen 41: 31-33.

Blick, T. and Harvey, M.S. (2011). Worldwide catalogues and species numbers of the arachnid orders (Arachnida). Arachnologische Mitteilungen 41: 41-43.

Harvey, M.S., Rix, M.G., Framenau, V.W., Hamilton, Z.R., Johnson, M.S., Teale, R.J., Humphreys, G. and Humphreys, W.F. (2011). Protecting the innocent: studying short-range endemic taxa enhances conservation outcomes. Invertebrate Systematics 25: 1-10.

Harvey, M.S. (2011). Notes on some Old World schizomids of the genera Ovozomus and Schizomus (Schizomida: Hubbardiidae). Records of the Western Australian Museum 26: 202-208. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2011_26(2)_202to208_HARVEY.pdf

Rix, M.G. and Harvey, M.S. (2011). Australian assassins, part I: A review of the Assassin Spiders (Araneae: Archaeidae) of mid-eastern Australia. Zookeys 123: 1-100. http://www.pensoft.net/journals/zookeys/article/1448/australian-assassins-part-i-a-review-of-the-assassin-spiders-araneae-archaeidae-of-mid-eastern-australia

Cooper, S.J.B., Harvey, M.S., Saint, K.M. and Main, B.Y. (2011). Deep phylogeographic structuring of populations of the trapdoor spider Moggridgea tingle (Migidae) from southwestern Australia: evidence for long-term refugia within refugia. Molecular Ecology 20: 3219-3236.

Rahmadi, C., Harvey, M.S. and Kojima, J. (2011).  The status of the whip spider subgenus Neocharon (Amblypygi: Charontidae) and the distribution of the genera Charon and Stygophrynus. Journal of Arachnology 39: 223-229. http://www.americanarachnology.org/JoA_free/JoA_v39_n2/arac-39-02-223.pdf

Harvey, M.S. (2011). Order Pseudoscorpiones de Geer, 1778. In: Zhang, Z.-Q. (ed.) Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 119-120.

2012

Rix, M.G. and Harvey, M.S. (2012). Phylogeny and historical biogeography of ancient assassin spiders (Araneae: Archaeidae) in the Australian mesic zone: evidence for Miocene speciation within Tertiary refugia. Molecular Phylogenetics and Evolution 62: 375-396.

Platnick, N.I., Abrahim, N., Álvarez-Padilla, F., Andriamalala, D., Baehr, B.C., Baert, L., Bonaldo, A.B., Brescovit, A.D., Chousou-Polydouri, N., Dupérré, N., Eichenberger, B., Fannes, W., Gaublomme, E., Gillespie, R.G., Grismado, C.J., Griswold, C.E., Harvey, M.S., Henrard, A., Hormiga, G., Izquierdo, M.A., Jocqué, R., Kranz-Baltensperger, Y., Kropf, C., Ott, R., Ramirez, M.J., Raven, R.J., Rheims, C.A., Ruiz, G.R.S., Santos, A.J., Saucedo, A., Sierwald, P., Szüts, T., Ubick, D. and Wang, X.-P. (2012). Tarsal organ morphology and the phylogeny of goblin spiders (Araneae, Oonopidae), with notes on basal genera. American Museum Novitates 3736: 1-52. http://digitallibrary.amnh.org/dspace/handle/2246/6160

Rix, M.G. and Harvey, M.S. (2012). Australian assassins, part II: A review of the new assassin spider genus Zephyrarchaea (Araneae, Archaeidae) from southern Australia. Zookeys 191: 1-62. http://www.pensoft.net/journals/zookeys/article/3070/australian-assassins-part-ii-a-review-of-the-new-assassin-spider-genus-zephyrarchaea-araneae-archaeidae-from-southern-au

Wojcieszek, J.M., Austin, P., Harvey, M.S. and Simmons, L.W. (2012). Micro-CT scanning provides insight into the functional morphology of millipede genitalia. Journal of Zoology, London 287: 91-95.

Fontaine, B., Achterberg, K. van, Alonso-Zarazaga, M.A, Araujo, R., Asche, M., Aspöck, H., Aspöck, Audisio, P., Aukema, B., Bailly, N., Balsamo, M, Bank, R.A., Belfiore, C., Bogdanowicz, W., Boxshall, G., Burckhardt, D., Chylarecki, P., Deharveng, L., Dubois, A., Enghoff, H., Fochetti, R., Fontaine, C., Gargominy, O., Gomez Lopez, M.S., Goujet, D., Harvey, M.S., Heller, K.-G., Helsdingen, P. van, Hoch, H., Jong, Y. De, Karsholt, O., Los, W., Magowski, W, Massard, J.A., McInnes, S.J., Mendes, L.F., Mey, E., Michelsen, V., Minelli, A., Nieto Nafrıa, J.M., van Nieukerken, E.J., Pape, T., De Prins, W., Ramos, M., Ricci, C., Roselaar, C., Rota, E., Segers, H, Timm, T., van Tol, J. and Bouchet, P. (2012). New species in the Old World: Europe as a frontier in biodiversity exploration, a test bed for 21st century taxonomy. PloS One 7: e36881. http://www.plosone.org/article/info%3Adoi%2F10.1371%2Fjournal.pone.0036881

Harvey, M.S. (2012). A new species of Synsphyronus (Pseudoscorpiones: Garypidae) from Western Australia. Records of the Western Australian Museum 27: 55-61. http://museum.wa.gov.au/sites/default/files/RecWAMuseum_2012_27(1)_055to061_HARVEY.pdf

Baehr, B.C., Harvey, M.S., Burger, M. and Thoma, M. (2012). The new Australasian goblin spider genus Prethopalpus (Araneae, Oonopidae). Bulletin of the American Museum of Natural History 763: 1-113. http://digitallibrary.amnh.org/dspace/handle/2246/6185

Volschenk, E.S., Harvey, M.S. and Prendini, L. (2012). A new species of Urodacus (Scorpiones: Urodacidae) from Western Australia. American Museum Novitates 3748: 1-18. http://digitallibrary.amnh.org/dspace/handle/2246/6181

Harvey, F.S.B., Framenau, V.W., Wojcieszek, J.M., Rix, M.G. and Harvey, M.S. (2012). Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia. Zootaxa 3383: 15-38. http://www.mapress.com/zootaxa/2012/f/z03383p038f.pdf

Sťáhlavský, F., Král, J., Harvey, M.S. and Haddad, C.R. (2012). The first cytogenetic characterisation of atemnids: pseudoscorpions with the highest chromosome numbers (Arachnida: Pseudoscorpiones). Cytogenetic and Genome Research 137: 22-30.

Rix, M.G. and Harvey, M.S. (2012). Australian assassins, part III: A review of the assassin spiders (Araneae, Archaeidae) of tropical north-eastern Queensland. Zookeys 218: 1-55. http://www.pensoft.net/journals/zookeys/article/3662/australian-assassins-part-iii-a-review-of-the-assassin-spiders-araneae-archaeidae-of-tropical-north-eastern-queensland

Harvey, M.S., Rahmadi, C., and Aland, K. (2012) The first record of whip spiders (Amblypygi) in Western Australia. Australasian Arachnology 84: 8-9.

Šťáhlavský, F., Boyer, S.L., Harvey, M.S. and Giribet, G. (2012). First cytogenetic study of a member of the harvestman family Pettalidae (Opiliones: Cyphophthalmi). Australian Journal of Entomology 51: 299-302.

Harvey, M.S., Ratnaweera, P.B. , Randeniya, P.V.  and Wijesinghe, M.R. (2012). A new species of the pseudoscorpion genus Megachernes (Pseudoscorpiones: Chernetidae) associated with a threatened Sri Lankan rainforest rodent, with a review of host associations of Megachernes. Journal of Natural History 46: 2519-2535.

2013

Car, C.A. and Harvey, M.S. (2013). A review of the Western Australian keeled millipede genus Boreohesperus (Diplopoda: Polydesmida: Paradoxosomatidae). Zookeys 290: 1–19. http://www.pensoft.net/journals/zookeys/article/5114/a-review-of-the-western-australian-keeled-millipede-genus-boreohesperus-diplopoda-polydesmida-paradoxosomatidae-

Zawierucha, K., Szymkowiak, P., Dabert, M. and Harvey, M.S. (2013). First record of the schizomid Stenochrus portoricensis (Schizomida: Hubbardiidae) in Poland, with DNA barcode data. Turkish Journal of Arachnology 37: 357–361. http://journals.tubitak.gov.tr/zoology/issues/zoo-13-37-3/zoo-37-3-11-1210-9.pdf

Baehr, B.C. and Harvey, M.S. (2013). The first goblin spiders of the genus Camptoscaphiella (Araneae: Oonopidae) from New Caledonia. Australian Journal of Entomology 52: 144–150.

Harms, D. and Harvey, M.S. (2013). Review of the cave-dwelling species of Pseudotyrannochthonius Beier (Arachnida: Pseudoscorpiones: Pseudotyrannochthoniidae) from mainland Australia, with description of two troglobitic species. Australian Journal of Entomology 52: 129–143.

Monod, L., Harvey, M.S. and Prendini, L. (2013). Stenotopic Hormurus Thorell, 1876 scorpions from the monsoon ecosystems of northern Australia, with a discussion on the evolution of burrowing behaviour in Hormuridae Laurie, 1896. Revue Suisse de Zoologie 120: 281–346.

Harvey, M.S. (2013). Order Pseudoscorpiones. Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness (Addenda 2013). Zootaxa 3703: 34–35. http://biotaxa.org/Zootaxa/article/view/zootaxa.3703.1.8/4262

Girişgin, A.O., Girişgin, O. and Harvey, M.S. (2013). The first record of pseudoscorpions in honeybee hives in Turkey. Turkish Journal of Veterinary and Animal Sciences 37: 608–610. http://journals.tubitak.gov.tr/veterinary/issues/vet-13-37-5/vet-37-5-21-1207-7.pdf

Nassirkhani, M. and Harvey, M.S. (2013). Description of the postlarval stages of Dactylochelifer gracilis Beier, Pseudoscorpiones: Cheliferidae. International Journal of Zoology 2013(782638): 1–15. http://www.hindawi.com/journals/ijz/2013/782638/

Baehr, B.C., Harvey, M.S., Smith, H.M. and Ott, R. (2013). The goblin spider genus Opopaea in Australia and the Pacific islands (Araneae: Oonopidae). Memoirs of the Queensland Museum – Nature 58: 107–338.

Harvey, M.S. and Baehr, B. (2013). Two new species of the goblin spider genus Cavisternum from tropical Australia (Araneae: Oonopidae). Memoirs of the Queensland Museum - Nature 58: 359–366.

Harvey, M.S. and Muchmore, W.B. (2013). The systematics of the pseudoscorpion family Ideoroncidae (Pseudoscorpiones, Neobisioidea) in the New World. Journal of Arachnology 41: 229–290.

Gunawardene, N.R., Taylor, C.K., Majer, J.D. and Harvey, M.S. (eds) (2013). The terrestrial invertebrate fauna of Barrow Island, Western Australia. Records of the Western Australian Museum, Supplement. Western Australian Museum: Perth. http://museum.wa.gov.au/store/museum-books/supplements/supplement-83-terrestrial-invertebrate-fauna-barrow-island

Humphreys, G., Alexander, J., Harvey, M.S. and Humphreys, W.F. (2013). The subterranean fauna of Barrow Island, north-western Australia: 10 years on. Records of the Western Australian Museum, Supplement 83: 145–158.

Car, C.A., Short, M., Huynh, C. and Harvey, M.S. (2013). The millipedes of Barrow Island, Western Australia (Diplopoda). Records of the Western Australian Museum, Supplement 83: 209–219.

Majer, J.D., Gunawardene, N.R., Taylor, C.K. and Harvey, M.S. (2013). A last word. Records of the Western Australian Museum, Supplement 83: 405–406.

Car, C.A., Wojcieszek, J.M. and Harvey, M.S. (2013). The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae) Part 1: Redefinition of the genus and redescriptions of existing species. Records of the Western Australian Museum 28: 83–118. http://museum.wa.gov.au/sites/default/files/WAMRecords_2013_28(2)_83to118_CARetal.pdf

Harvey, M.S. (2014). A review and redescription of the cosmopolitan pseudoscorpion Chelifer cancroides (Pseudoscorpiones: Cheliferidae). Journal of Arachnology 42: 86–104

Harvey, M.S. and Du Preez, G. (2014). A new troglobitic ideoroncid pseudoscorpion (Pseudoscorpiones: Ideoroncidae) from southern Africa. Journal of Arachnology 42: 105–110.