Dr Mark Harvey

Senior Curator of Arachnology

Terrestrial Zoology

B.Sc., Ph.D.

"Dr Mark Harvey working at a microscope"


+61 (8) 9212 3737 mark.harvey@museum.wa.gov.au


  • Systematics of arachnids and myriapods, including their phylogenetic history and evolutionary relationships.
  • Taxonomy of arachnids and myriapods, and especially the documentation and description of new species.


  • Researches the taxonomy and systematics of arachnids and myriapods through the collections.
  • Develops the Museum’s collections through targeted fieldwork and collaborations with other organisations, including government, universities and industry.

Services (to other bodies):

  • Represents the Museum on committees and panels providing advice to government.


Exhibitions and Public Programs

  • Provides identifications and advice for environmental consultants, industry and government agencies.
  • Regularly presents talks and provides media coverage of work on arachnids and myriapods.
  • Co-editor of the Records of the Western Australian Museum; Associate Editor of the Journal of Arachnology and Invertebrate Systematics. Member of the editorial board of several other scholarly journals.
  • Co-supervisor to several post-graduate students at local and international universities.
  • Vice-President of the International Commission on Zoological Nomenclature: http://iczn.org/content/dr-mark-s-harvey

Selected Publications

Mark’s full publication list can be found here



Harvey, M.S. and Evenhuis, N.L. (2020). Revised authorship and dates of some genus-group names in Arachnida proposed without type species after 1930. Arachnology 18: 623–628.

Harvey, M.S. and Cullen, K.L. (2020). A remarkable new troglobitic species of Parobisium (Pseudoscorpiones: Neobisiidae) from California. Arachnology 18: 591–596.

Abrams, K.M., Huey, J.A., Hillyer, M.J., Didham, R.K. and Harvey, M.S. (2020). A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara. Zootaxa 4864: 1–75.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2020). The open-holed trapdoor spiders (Mygalomorphae: Anamidae: Namea) of Australia’s D’Aguilar Range: revealing an unexpected subtropical hotspot of rainforest diversity. Zootaxa 4861: 71–91.

Castalanelli, M.A., Framenau, V.W., Huey, J.A. and Harvey, M.S. (2020). New species of the open-holed trapdoor spider genus Aname (Araneae: Mygalomorphae: Anamidae) from arid Western Australia. Journal of Arachnology 48: 169–213.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2020). First phylogenetic assessment and taxonomic synopsis of the open-holed trapdoor spider genus Namea (Mygalomorphae: Anamidae): a highly diverse mygalomorph lineage from Australia’s tropical eastern rainforests. Invertebrate Systematics 34: 679–726.

Harvey, M.S. (2020). Synsphyronus platnicki sp. nov.: first Synsphyronus (Pseudoscorpiones: Garypidae) from New Caledonia. Arachnology 18: 468–472.

Harvey, M.S., Hillyer, M.A., Carvajal, J.I. and Huey, J.A. (2020). Supralittoral pseudoscorpions of the genus Garypus (Pseudoscorpiones: Garypidae) from the Indo-West Pacific region, with a review of the subfamily classification of Garypidae. Invertebrate Systematics 34: 34—87.

Abrams, K.M., Huey, J.A., Hillyer, M.A., Humphreys, W.F., Didham, R.K. and Harvey, M.S. (2019). Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution. 139 (106532): 1–12.

Benavides, L.R., Cosgrove, J.G., Harvey, M.S. and Giribet, G. (2019). Phylogenomic interrogation resolves the backbone of the Pseudoscorpiones Tree of Life. Molecular Phylogenetics and Evolution. 139 (106509): 1–14.

Huey, J.A., Rix, M.G., Wilson, J.D., Hillyer, M.J. and Harvey, M.S. (2019). Open-holed trapdoor spiders of the genus Teyl (Mygalomorphae: Nemesiidae: Anamini) from Western Australia’s Pilbara bioregion: a new species and expanded phylogenetic assessment. Zootaxa 4674: 349–362.

Harms, D., Roberts, J.D. and Harvey, M.S. (2019). Climate variability impacts on diversification processes in a biodiversity hotspot: a phylogeography of ancient pseudoscorpions in southwestern Australia. Zoological Journal of the Linnean Society 186: 934–949.

Huey, J.A., Hillyer, M.J. and Harvey, M.S. (2019). Phylogenetic relationships and biogeographic history of the Australian trapdoor spider genus Conothele (Araneae: Mygalomorphae: Halonoproctidae): diversification into arid habitats in an otherwise tropical radiation. Invertebrate Systematics 33: 628–643.

Ott, R., Ubick, D., Bonaldo, A.B., Brescovit, A.D. and Harvey, M.S. (2019). A revision of the New World goblin spider genus Cinetomorpha Simon, 1892 revalidated from Gamasomorpha Karsch, 1881 (Araneae, Oonopidae, Oonopinae). Zootaxa 4641: 1–152.

Car, C.A., Harvey, M.S. and Huey, J.A. (2019). The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 3: species of the Pilbara bioregion of Western Australia. Zootaxa 4617: 1–71.

Gibson, L., Humphreys, W.F., Harvey, M.S., Hyder, B. and Winzer, A. (2019). Shedding light on the hidden world of subterranean fauna: a transdisciplinary research approach. Science of the Total Environment 684: 381–389.

Rix, M.G., Wilson, J.D., Rix, A.G., Wojcieszek, A.M., Huey, J.A. and Harvey, M.S. (2019). Population demography and biology of a giant spiny trapdoor spider (Idiopidae: Euoplos) from inland Queensland: developing a ‘slow science’ study system to address a conservation crisis. Austral Entomology 58: 282–297.

Novák, J. and Harvey, M.S. (2019). The status of the pseudoscorpion genus Indogarypus Beier, 1957 with new descriptions of three species of Geogarypus Chamberlin, 1930 (Arachnida: Pseudoscorpiones: Geogarypidae). Journal of Arachnology 47: 124–131.

Johnson, J., Romero-Ortiz, C., Mathew, A.V., Sebastian, P.A., Joseph, M.M. and Harvey, M.S. (2019). A review of the pseudoscorpion genus Metawithius (Pseudoscorpiones: Withiidae) from the Indian subcontinent. Journal of Arachnology 47: 84–94.

Rix, M.G., Wilson, J.D. and Harvey, M.S. (2019). A revision of the white-headed spiny trapdoor spiders of the genus Euoplos (Mygalomorphae: Idiopidae: Arbanitinae): a remarkable lineage of rare mygalomorph spiders from the south-western Australian biodiversity hotspot. Journal of Arachnology 47: 63–76.

Harvey, M.S. and Mahnert, V. (2019). The status of the Central American pseudoscorpion genus Coprochernes Beier, a new synonym of Neoallochernes Hoff (Pseudoscorpiones: Chernetidae). Arachnology 18: 7–13.

Romero-Ortiz, C. and Harvey, M.S. (2019). The pseudoscorpion genus Verrucachernes (Pseudoscorpiones: Chernetidae) in the Indian region. Zootaxa 4569: 337–346.

Kropf, C., Blick, T., Brescovit, A.D., Chatzaki, M., Dupérré, N., Gloor, D., Haddad, C.R., Harvey, M.S., Jäger, P., Marusik, Y.M., Ono, H., Rheims, C.A. and Nentwig, W. (2019). How not to delimit taxa: a critique on a recently proposed “pragmatic classification” of jumping spiders (Arthropoda: Arachnida: Araneae: Salticidae). Zootaxa 4545: 444–446.

Framenau, V.W., Hamilton, Z.R., Finston, T., Humphreys, G., Abrams, K.M., Huey, J.A. and Harvey, M.S. (2018). Molecular and morphological characterization of new species of hypogean Paradraculoides (Schizomida: Hubbardiidae) from the arid Pilbara bioregion of Western Australia. Journal of Arachnology 46: 507–537.

Rix, M.G., Raven, R.J. and Harvey, M.S. (2018). Systematics of the giant spiny trapdoor spiders of the genus Gaius Rainbow (Mygalomorphae: Idiopidae: Aganippini): documenting an iconic lineage of the Western Australian inland arid zone. Journal of Arachnology 46: 438–472.

Chamberland, L., McHugh, A., Kechejian, S., Binford, G., Bond, J.E., Coddington, J.A., Dolman, G., Hamilton, C.A., Harvey, M.S., Kuntner, M. and Agnarsson, I. (2018). From Gondwana to GAARlandia: evolutionary history and biogeography of ogre‐faced spiders (Deinopis). Journal of Biogeography 45: 2442–2457.

Sato, S., Buckman-Young, R.S., Harvey, M.S. and Giribet, G. (2018). Cryptic speciation in a biodiversity hotspot: multilocus molecular data reveal new velvet worm species from Western Australia (Onychophora: Peripatopsidae: Kumbadjena). Invertebrate Systematics 32: 1249–1264.

Harrison, S.E., Rix, M.G., Harvey, M.S. and Austin, A.D. (2018). Systematics of the Australian spiny trapdoor spiders of the genus Blakistonia Hogg (Araneae: Idiopidae). Zootaxa 4518: 1–76.

Harvey, M.S., Hillyer, M.A., Main, B.Y., Moulds, T.A., Raven, R.J., Rix, M.G., Vink, C.J. and Huey, J.A. (2018). Phylogenetic relationships of the Australasian open-holed trapdoor spiders (Araneae: Mygalomorphae: Nemesiidae: Anaminae): multi-locus molecular analyses resolve the generic classification of a highly diverse lineage. Zoological Journal of the Linnean Society 184: 407–452. doi.org/10.1093/zoolinnean/zlx111.

Majer, J.D., Harvey, M.S., Humphreys, W.F., Davis, J.A. and Yen, A.L. (2018). Terrestrial and inland-water invertebrates of the Australian arid zone. In: H. Lambers, On the Ecology of Australia’s Arid Zone: 181–213. (Springer International Publishing).

Harvey, M.S. (2018). Balgachernes occultus, a new genus and species of pseudoscorpion (Pseudoscorpiones: Chernetidae) associated with balga (Xanthorrhoea preissii) in south-western Australia, with remarks on Austrochernes and Troglochernes. Records of the Western Australian Museum 33: 115–130.

Rix, M.G., Raven, R.J., Austin, A.D., Cooper, S.J.B. and Harvey, M.S. (2018). Systematics of the spiny trapdoor spider genus Bungulla (Mygalomorphae: Idiopidae): revealing a remarkable radiation of mygalomorph spiders from the Western Australian arid zone. Journal of Arachnology 46: 249–344.

Harvey, M.S., Gardini, G. and Novák, J. (2018). Case 3736 – Garypus latus Hansen, 1884 (currently Larca lata; Arachnida, Pseudoscorpiones): proposed precedence over Garypus hungaricus Tömösváry, 1882. Bulletin of Zoological Nomenclature 75: 16–20.

Santibañez-Lopez, C.E., Ontano, A.Z., Harvey, M.S. and Sharma, P.P. (2018). Transcriptomic analysis of pseudoscorpion venom reveals a unique cocktail dominated by enzymes and protease inhibitors. Toxins 10(207): 1–12.

Taylor, G.S., Braby, M.F., Moir, M.L., Harvey, M.S., Sands, D.P.A., New, T.R., Kitching, R.L., Hogendoorn, K., Glatz, R.V., Andren, M., Cook, J.M., McQuillan, P.B., Henry, S.C., Valenzuela, I. and Weinstein, P. (2018). A strategic national approach for improving the conservation management of insects and allied invertebrates in Australia. Austral Entomology 57: 124–149.

Rix, M.G., Huey, J.A., Cooper, S.J.B., Austin, A.D. and Harvey, M.S. (2018). Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia. ZooKeys 756: 1–121.

Harvey, M.S. (2018). Two new species of the pseudoscorpion genus Cybella (Pseudoscorpiones: Feaellidae) from Malaysian caves. Zoologischer Anzeiger 273: 124–132.

Harvey, M.S., Cosgrove, J.G., Harms, D., Selden, P.A., Shih, C. and Wang, C.-C. (2018). The oldest chthonioid pseudoscorpion (Arachnida: Pseudoscorpiones: Chthonioidea: Chthoniidae): a new genus and species from mid-Cretaceous Burmese amber. Zoologischer Anzeiger 273: 102–111.

Rix, M.G., Main, B.Y., Raven, R.J. and Harvey, M.S. (2018). Systematics of the spiny trapdoor spiders of the genus Eucanippe (Mygalomorphae: Idiopidae: Aganippini) from south-western Australia: documenting a poorly-known lineage from Australia's biodiversity hotspot. Journal of Arachnology 46: 133–154.

Novák, J. and Harvey, M.S. (2018). New species and records of the pseudoscorpion genus Geogarypus (Pseudoscorpiones: Geogarypidae) from India, Sri Lanka and New Guinea. Zootaxa 4394: 417–427.

Thomson, S.A., Pyle, R.L., Ahyong, S., Alonso-Zarazaga, M., Ammirati, J., Araya, J.F., Ascher, J.S., Audisio, T.L., Azevedo-Santos, V.M., Bailly, N., Baker, W.J., Balke, M., Barclay, M.V.L., Barrett, R.L., Benine, R.C., Bickerstaff, J.R.M., Bouchard, P., Bour, R., Bourgoin, T., Boyko, C.B., Breure, A.S.H., Brothers, D.J., Byng, J.W., Campbell, D., P.Ceriaco, L.M., Cernak, I., Cerretti, P., Chang, C.-H., Cho, S., Copus, J.M., Costello, M.J., Cseh, A., Csuzdi, C., Culham, A., D’Elia, G., d’Acoz, C.d.U., Daneliya, M.E., Dekker, R., Dickinson, E.C., Dickinson, T.A., Dijk, P.P.v., Dijkstra, K.-D.B., Dima, B., Dmitriev, D.A., Duistermaat, L., Dumbacher, J.P., Eiserhardt, W.L., Ekrem, T., Evenhuis, N.L., Faille, A., Fernandez-Triana, J.L., Fiesler, E., Fishbein, M., Fordham, B.G., Freitas, A.V.L., Friol, N.R., Fritz, U., Froslev, T., Funk, V.A., Gaimari, S.D., Garbino, G.S.T., S.Garraffoni, A.R., Geml, J., Gill, A.C., Gray, A., Grazziotin, F.G., Greenslade, P., Gutierrez, E.E., Harvey, M.S., Hazevoet, C.J., He, K., He, X., Helfer, S., Helgen, K.M., Heteren, A.H.v., Garcia, F.H., Holstein, N., Horvath, M.K., Hovenkamp, P.H., Hwang, W.S., Hyvonen, J., Islam, M.B., Iverson, J.B., Ivie, M.A., Jaafar, Z., Jackson, M.D., Jayat, J.P., Johnson, N.F., Kaiser, H., Klitgard, B.B., Knapp, D.G., Kojima, J.-i., Koljalg, U., Kontschan, J., Krell, F.-T., Krisai-Greilhuber, I., Kullander, S., Latella, L., Lattke, J.E., Lencioni, V., Lewis, G.P., Lhano, M.G., Lujan, N.K., Luksenburg, J.A., Mariaux, J., Marinho-Filho, J., Marshall, C.J., Mate, J.F., McDonough, M.M., Michel, E., Miranda, V.F.O., Mitroiu, M.-D., Molinari, J., Monks, S., Moore, A.J., Moratelli, R., Muranyi, D., Nakano, T., Nikolaeva, S., Noyes, J., Ohl, M., Oleas, N.H., Orrell, T., Pall-Gergely, B., Pape, T., Papp, V., Parenti, L.R., Patterson, D., Pavlinov, I.Y., Pine, R.H., Poczai, P., Prado, J., Prathapan, D., Rabeler, R.K., Randall, J.E., Rheindt, F.E., Rhodin, A.G.J., Rodriguez, S.M., Rogers, D.C., Roque, F.d.O., Rowe, K.C., Ruedas, L.A., Salazar-Bravo, J., Salvador, R.B., Sangster, G., Sarmiento, C.E., S.Schigel, D., Schmidt, S., Schueler, F.W., Segers, H., Snow, N., Souza-Dias, P.G.B., Stals, R., Stenroos, S., Stone, R.D., Sturm, C.F., Štys, P., Teta, P., Thomas, D.C., Timm, R.M., Tindall, B.J., Todd, J.A., Triebel, D., Valdecasas, A.G., Vizzini, A., Vorontsova, M.S., Vos, J.M.d., Wagner, P., Watling, L., Weakley, A., Welter-Schultes, F., Whitmore, D., Wilding, N., Will, K., Williams, J., Wilson, K., Winston, J.E., Wuster, W., Yanega, D., Yeates, D.K., Zaher, H., Zhang, G., Zhang, Z.-Q. and Zhou, H.-Z. (2018). Taxonomy based on science is necessary for global conservation. PloS Biology 16 (3): e2005075.

Stephen, C.D.R. and Harvey, M.S. (2018). William B. Muchmore (1920–2017): his taxonomic contributions and complete bibliography. Journal of Cave and Karst Studies 80: 39–48.

Schmidt, A.R., Kaulfuss, U., Bannister, J.M., Baranov, V., Beimforde, C., Bleile, N., Borkent, A., Busch, A., Conran, J.G., Engel, M.S., Harvey, M., Kennedy, E.M., Kerr, P.H., Kettunen, E., Kiecksee, A.P., Lengeling, F., Lindqvist, J.K., Maraun, M., Mildenhall, D.C., Perrichot, V., Rikkinen, J., Sadowski, E.-M., Seyfullah, L.J., Stebner, F., Szwedo, J., Ulbrich, P. and Lee, D.E. (2018). Amber inclusions from New Zealand. Gondwana Research 56: 135–146.

Sharma, P.P., Baker, C.M., Cosgrove, J.G., Johnson, J.E., Oberski, J.T., Raven, R.J., Harvey, M.S., Boyer, S.L. and Giribet, G. (2018). A revised dated phylogeny of scorpions: Phylogenomic support for ancient divergence of the temperate Gondwanan family Bothriuridae. Molecular Phylogenetics and Evolution 122: 37–45.

Rix, M.G., Bain, K., Main, B.Y., Raven, R.J., Austin, A.D., Cooper, S.J.B. and Harvey, M.S. (2017). The  spiny trapdoor spiders of the genus Cataxia (Mygalomorphae: Idiopidae) from south-western Australia: documenting a threatened fauna in a sky-island landscape. Journal of Arachnology 45: 395–423.

Clouse, R.M., Branstetter, M.G., Buenavente, P., Crowley, L.M., Czekanski-Moir, J., Emmanuel, M., General, D., Giribet, G., Harvey, M.S., Janies, D.A., Mohagan, A.B., Mohagan, D.P., Sharma, P.P. and Wheeler, W.C. (2017). First global molecular phylogeny and biogeographical analysis of two arachnid orders (Schizomida and Uropygi) supports a tropical Pangean origin and mid-Cretaceous diversification. Journal of Biogeography 44: 2660–2672.

Rix, M.G., Raven, R.J., Main, B.Y., Harrison, S.E., Austin, A.D., Cooper, S.J.B. and Harvey, M.S. (2017). The Australasian spiny trapdoor spiders of the family Idiopidae (Mygalomorphae: Arbanitinae): a relimitation and revision at the generic level. Invertebrate Systematics 31: 566–634.

Harrison, S.E., Harvey, M.S., Cooper, S.J.B., Austin, A.D. and Rix, M.G. (2017). Across the Indian Ocean: A remarkable example of trans-oceanic dispersal in an austral mygalomorph spider. PLoS One. 12(8) e0180139: 1–16.

Castalanelli, M.A., Huey, J.A., Hillyer, M.J. and Harvey, M.S. (2017). Molecular and morphological evidence for a new genus of small trapdoor spiders from arid Western Australia (Araneae: Mygalomorphae: Nemesiidae: Anaminae). Invertebrate Systematics 31: 492–505.

Smith, H.M., Harvey, M.S., Agnarsson, I. and Anderson, G.J. (2017). Notes on the ant-mimic genus Anatea Berland (Araneae: Theridiidae) and two new species from tropical Australia. Records of the Australian Museum 69: 1–13.

Car, C.A. and Harvey, M.S. (2017). New species of Boreohesperus (Diplopoda, Polydesmida, Paradoxosomatidae) from north-western Australia. European Journal of Taxonomy 320: 1–11.

Gao, Z., Zhang, F. and Harvey, M.S. (2017). A modified definition of the genus Haplochernes (Pseudoscorpiones: Chernetidae), with a new species from Hainan Island. Journal of Arachnology 45: 112–122.

Rix, M.G., Huey, J.A., Main, B.Y., Waldock, J.M., Harrison, S.E., Comer, S., Austin, A.D. and Harvey, M.S. (2017). Where have all the spiders gone? Highlighting the decline of a poorly known invertebrate fauna in the agricultural and arid zones of southern Australia. Austral Entomology 56: 14–22.

Rix, M.G., Cooper, S.J.B., Meusemann, K., Klopfstein, S., Harrison, S.E., Harvey, M.S. and Austin, A.D. (2017). Post-Eocene climate change across continental Australia and the diversification of Australasian spiny trapdoor spiders (Idiopidae: Arbanitinae) Molecular Phylogenetics and Evolution 109: 302–320.

Harvey, M.S., Rix, M.G., Harms, D., Giribet, G., Vink, C.J. and Walter, D.E. (2017). The biogeography of Australasian arachnids. In: M. Ebach (ed.), Handbook on Australasian Biogeography. (CRC Press / Taylor and Francis: Boca Raton).

Harvey, M.S. and Bosmans, R. (2016). Order Pseudoscorpiones, pseudoscorpions. In: D. J. Sparrow and E. John, An introduction to the wildlife of Cyprus: 521–524. (Terra Cypria: Lemesos).

Harvey, M.S., Andrade, R. and Pinto-da-Rocha, R. (2016). The first New World species of the pseudoscorpion family Feaellidae (Pseudoscorpiones: Feaelloidea), from the Brazilian Atlantic Forest. Journal of Arachnology 44: 237–234.

Harrison, S.E., Rix, M.G., Harvey, M.S. and Austin, A.D. (2016). An African mygalomorph lineage in temperate Australia: the trapdoor spider genus Moggridgea (Araneae: Migidae) on Kangaroo Island, South Australia. Austral Entomology 55: 208–216.

Downes, M.F. and Harvey, M.S. (2016). The spider Oecobius concinnus Simon (Araneae: Oecobiidae) in nests of the ant Polyrhachis australis Mayr (Hymenoptera: Formicidae) in Townsville, Queensland: a new distribution record. Australian Entomologist 43: 31–34.

Harvey, M.S., Huey, J.A., Hillyer, M.A., McIntyre, E. and Giribet, G. (2016) The first troglobitic species of Gymnobisiidae (Pseudoscorpiones, Neobisioidea), from Table Mountain (Western Cape Province, South Africa) and its phylogenetic position. Invertebrate Systematics 30: 75–85.

Harvey, M.S. (2016). The systematics of the pseudoscorpion family Ideoroncidae (Pseudoscorpiones, Neobisioidea) in the Asian region. Journal of Arachnology 44: 272–329.

Cosgrove, J.G., Agnarsson, I., Harvey, M.S. and Binford, G.J. (2016). Pseudoscorpion diversity and distribution in the West Indies: sequence data confirm single island endemism for some clades, but not others. Journal of Arachnology 44: 257–271.

Harvey, M.S., Abrams, K.M., Beavis, A.S., Hillyer, M.J. and Huey, J.A. (2016). Pseudoscorpions of the family Feaellidae (Pseudoscorpiones: Feaelloidea) from the Pilbara region of Western Australia show extreme short-range endemism. Invertebrate Systematics 30: 491–508.

Giribet, G., Boyer, S.L., Baker, C., Fernández, R., Sharma, P., De Bivort, B.L., Daniels, S.R., Harvey, M.S., Neethling, J.A. and Griswold, C.E. (2016). A molecular phylogeny of the temperate Gondwanan family Pettalidae (Arachnida, Opiliones, Cyphophthalmi) with biogeographic and taxonomic implications. Zoological Journal of the Linnean Society 178: 523–545.


Selected publications (pre-2016)

Harvey, M.S., Main, B.Y., Rix, M.G. and Cooper, S.J.B. (2015). Refugia within refugia: in situ speciation and conservation of threatened Bertmainius (Araneae: Migidae), a new genus of relictual trapdoor spiders endemic to the mesic zone of south-western Australia. Invertebrate Systematics 29: 511–553.

Rix, M.G., Edwards, D.L., Byrne, M., Harvey, M.S., Joseph, L. and Roberts, J.D. (2015). Biogeography and speciation of terrestrial fauna in the south-western Australian biodiversity hotspot. Biological Reviews 90: 762–793.

Rix, M.G. and Harvey, M.S. (2012). Phylogeny and historical biogeography of ancient assassin spiders (Araneae: Archaeidae) in the Australian mesic zone: evidence for Miocene speciation within Tertiary refugia. Molecular Phylogenetics and Evolution 62: 375–396.

Harvey, M.S., Rix, M.G., Framenau, V.W., Hamilton, Z.R., Johnson, M.S., Teale, R.J., Humphreys, G. and Humphreys, W.F. (2011). Protecting the innocent: studying short-range endemic taxa enhances conservation outcomes. Invertebrate Systematics 25: 1–10.

Murienne, J., Harvey, M.S. and Giribet, G. (2008). First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution 49: 170–184.