Atemnidae Kishida, 1929

Family

Oratemnus sp. from Australia  (Image: K. Edward)Paratemnoides sp. from Australia  (Image: K. Edward)

The Atemnidae were created by Kishida (1929b) and Chamberlin (1931) for Atemnus, and augmented with additional genera by Beier (1932) and later workers. The Miratemninae were added by Beier (1932) for Miratemnus. The miratemnines were considered a separate family by Dumitresco and Orghidan (1970), but it was returned as a subfamily of Atemnidae by Harvey (1992) which was supported by Klausen (2005).

Composition

The Atemnidae are divided into two subfamilies, Atemninae and Miratemninae. The Atemninae contain 15 Recent genera, and a single genus, Progonatemnus, from Eocene Baltic Amber. The Miratemninae contain six genera. Many atemnid genera appear to be poorly defined and significant changes will be necessary to create a more workable classification (Klausen 2005).

Distribution

Atemnines are found in most regions of the world, whereas miratemnines are more restricted, being found in Africa, southern Europe, central Asia, North America and South America.

Distribution Count
2
Afghanistan 6
Angola 2
Argentina 1
Austria 1
Australia 5
Azerbaijan 3
Barbados 1
Bulgaria 3
Burundi 1
Brazil 4
Bhutan 3
Democratic Republic of Congo (Zaire) 17
Congo 8
Switzerland 1
Ivory Coast 9
Cameroon 12
China 6
Colombia 1
Costa Rica 1
Cuba 1
Cape Verde 1
Cyprus 1
Dominican Republic 1
Algeria 3
Ecuador 3
Egypt 1
Spain 3
Ethiopia 5
Micronesia 4
France 1
Gabon 4
Georgia 1
Ghana 5
Guinea 5
Equatorial Guinea 6
Greece 2
Guatemala 2
Guam 2
Guinea-Bissau 5
Guyana 3
Croatia 1
Haiti 1
Hungary 1
Indonesia 22
Israel 1
India 30
Iran 3
Italy 1
Jamaica 2
Japan 4
Kenya 19
Kyrgyzstan 3
Cambodia 4
Saint Kitts and Nevis 2
Kazakhstan 3
Laos 4
Sri Lanka 12
Libya 1
Morocco 1
Madagascar 5
Marshall Islands 3
Macedonia 1
Myanmar 6
Mongolia 3
Northern Mariana Islands 4
Malawi 1
Mexico 2
Malaysia 10
Mozambique 1
Namibia 6
New Caledonia 1
Nigeria 2
Nepal 2
Panama 2
Peru 1
French Polynesia 2
Papua New Guinea 13
Philippines 13
Pakistan 1
Puerto Rico 1
Portugal 1
Palau 2
Paraguay 1
Romania 2
Rwanda 1
Solomon Islands 8
Seychelles 3
Sudan 3
Singapore 1
Slovakia 4
Sierra Leone 1
Senegal 1
Somalia 6
Sao Tome and Principe 4
Syria 2
Chad 4
French Southern Territories 1
Togo 2
Thailand 12
Tajikistan 3
Turkmenistan 3
Tunisia 2
Turkey 5
Trinidad and Tobago 1
Tanzania 11
Ukraine 1
Uganda 9
United States 2
Uzbekistan 3
Saint Vincent and the Grenadines 1
Venezuela 1
U.S. Virgin Islands 1
Vietnam 11
Vanuatu 2
Samoa 2
Yemen 1
Mayotte 1
South Africa 14
Zimbabwe 5

Important References

Klausen, F.E. (2005). The male genitalia of the family Atemnidae (Pseudoscorpiones). Journal of Arachnology 33: 641-662.

References

Pessigini Navás, 1925: 109. Atemnidae Kishida, 1929b: 124; Chamberlin, 1931a: 243-244; Beier, 1932b: 548; Beier, 1932e: 20; Beier, 1932g: 186; Roewer, 1937: 279; Hoff, 1956c: 4; Morikawa, 1960: 136; Beier, 1963b: 245; Murthy and Ananthakrishnan, 1977: 124; Muchmore, 1982a: 100; Harvey, 1985b: 128; Harvey, 1991a: 451; Harvey, 1992c: 1430-1431; Tooren, 2008: 435.